The famous grizzly bear mother, a longtime icon of the Greater Yellowstone Ecosystem, has died after being struck by a vehicle in Wyoming.

Grizzly bear 399 was killed on Tuesday after a vehicle hit it on U.S. Highway 26/89 in Snake River Canyon, south of Jackson, Wyoming. The bear’s identity was confirmed through ear tags and a microchip.

Collision was purely accidental and there was no indication of any wrongdoing, Lincoln County sheriff’s official says.